The website owner is responsible and that is who Getty chase for the money.
I would send a letter explaining why they came to be used, state you were not aware they were coyrighted and you made no profit from them, you have now removed them, apologise, ask why a take-down notice was not issued as per DMCA and that you dont recognise their invoice as valid or legal.
Read the sitepoint thread in full
Then ignore absolutely anything else they send.
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